Familiar faces include EastEnders' Phil Daniels, who previously appeared in season 1 as fisherman Marvellous Harris, and Gavin & Stacey star Melanie Walters, whose role has yet to be revealed.
Elsewhere, Humphrey (Kris Marshall) and Martha (Sally Bretton) "launch into married life as they search for a new home, while Humphrey wrestles with an impossible decision that may change the lives of everyone at the station forever".
But it remains to be seen whether they will continue to foster or even consider adoption – especially given just how painful it was to say goodbye to Rosie last season when she returned to her biological mum.
The guest cast for season 4 also includes Susan George (Straw Dogs), Colin McFarlane (The War Between the Land and the Sea), Michael Cochrane (The Archers), Vincent Franklin (The Thick of It), Alex Macqueen (The Inbetweeners) and Tristan Sturrock (Poldark).
Rounding it out are Andrea Lowe (Sherwood), Rachel Cassidy (Downton Abbey), Emily Stott (Foundation), Rina Mahoney (Happy Valley), Susan Penhaligon (A Fine Romance), James Clay (MobLand), Kenneth Collard (Cuckoo) and Narinda Samra (Ackley Bridge).
As for other potential storylines, after an initial health scare last season, Anne was given the all-clear – but will her new lease of life see her launch herself back into the dating scene?
Meanwhile, there still appears to be unfinished business between Esther and Archie, despite the detective continually pushing him away.
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